Great Red Spinners

This is by far the most important species of Isonychia. Many angling books once split its credit with the species Isonychia sadleri and Isonychia harperi, but entomologists have since discovered that those are just variations of this abundant species.

See the main Isonychia page for more about these intriguing mayflies.

This species (or rather group of subspecies (Subspecies: Entomologists sometimes further divide a species into distinct groups called subspecies, which have two lower-case words on the end of their scientific name instead of one. The latter is the sub-species name. For example, Maccaffertium mexicanum mexicanum and Maccaffertium mexicanum integrum are two different subspecies of Maccaffertium mexicanum.)), together with Drunella doddsii, make up the famous Western Green Drake hatches. They are widespread throughout the vast Western region and can be abundant enough in many locations to provide world class angling.

It hasn't been all that many years since Western traditions and entomological "facts on the ground" began to influence the angler's lexicon heavily dominated by Eastern writers. Their initial reporting after visiting the region first popularized the phrase "Rocky Mountains answer to the popular Green Drakes of the East". This led to a false impression that lingers to this day. The reality is these giants of their family have abundant populations all over the West with no counterpart in the East, and the West does have abundant hatches of comparable Ephemeridae. The Western tradition of naming outsized Mayflies "Drakes" is the reason for what many consider a misnomer by giving it the same common name as the legendary Ephemerid of the East and surely contributed to confusion for anglers unconcerned with such subtleties.

Timpanoga hecuba is not abundant enough, and its emergence not concentrated enough to provide great hatches, but where it is locally abundant it creates fishable action because of its large size. This species seems subject to substantial fluctuations in population densities, possibly in relation to the amount of silted habitat they prefer. When silt builds up in drought years, their numbers appear to increase. It is the largest species in the Ephemerellidae family, often rivaling Drunella grandis (Western Green Drake) in length but appearing even stouter due to its dramatic lateral (Lateral: To the side.) abdominal spines. It contains two subspecies (Subspecies: Entomologists sometimes further divide a species into distinct groups called subspecies, which have two lower-case words on the end of their scientific name instead of one. The latter is the sub-species name. For example, Maccaffertium mexicanum mexicanum and Maccaffertium mexicanum integrum are two different subspecies of Maccaffertium mexicanum.). See the Timpanoga genus hatch page for details.

There seems to be no preferred common name and anglers call them many things. Great Red Quill and Western Red Drake seem to fit best, though many refer to them as Giant Dark Hendricksons. Many fly shop reports have recently started to refer to them by their Latin species name. A fly shop serving Yellowstone out of Gardiner, Montana calls them Drake Mackerels in their hatch chart.

In the East and Midwest this is one of the most important hatches of the Spring. They are large flies which emerge sporadically, making for long days of good fishing.

This species contains the two classic Eastern hatches formerly known as Stenonema vicarium and Stenonema fuscum, the "March Brown" and "Gray Fox." Entomologists have discovered that these mayflies belong to the same species, but they still display differences in appearance which the trout notice easily. Anglers should be prepared to imitate both types.

This species together with the Drunella grandis sub-species make up the Western Green Drake hatch. Besides being smaller, the adults are difficult to tell apart from it's larger siblings; but D. doddsi nymphs have a few peculiar traits that set them apart. D. doddsi looks much thicker in the thorax (Thorax: The thorax is the middle part of an insect's body, in between the abdomen and the head, and to which the legs and wings are attached.), has a flat frontal head margin and a unique oval disk-like ring of hairs on its ventral (Ventral: Toward or on the bottom.) surface. However, There are very few differences between the habits of these two species, and they are almost always discussed together in fly fishing books, so for many of the characteristics of doddsii, refer to the Drunella grandis page.

The Brown Drakes are a favorite hatch of many in the Midwest, and they make a good showing on localized waters across the country. They are usually the first in a series of big drakes which bring large trout to the surface at twilight and into the early hours of the night.